Shadow Enhancers as a Source of Evolutionary Novelty

Science  05 Sep 2008:
Vol. 321, Issue 5894, pp. 1314
DOI: 10.1126/science.1160631

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Some developmentally important genes can be regulated via two enhancers, one located nearby and the other, a "shadow" enhancer, 10 to 20 kilobases away.

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